Myroxylon balsamum
(L.) Harms
Fabaceae
This species has at times been treated as two distinct species (Myroxylon balsamum (L.) Harms and Myroxylon pereirae (Harms) Klotzsch), but is now more commonly treated as two varieties of the same species, a treatment we are following here. This record is for the main species (commonly known as Tolu Balsam), see Myroxylon balsamum pereirae for the form commonly known as Peru Balsam.
Myrospermum toluiferum DC.
Myroxylon toluiferum A.Rich.
Toluifera balsamum L.
Common Name: Tolu Balsam
General Information
Myroxylon balsamum is a slow-growing, emergent tree of the rainforest that can reach a height of 45 metres but is usually smaller[
337- Title
- Tropical Tree Seed Manual
- Publication
-
- Author
- Vosso. J. Editor.
- Website
- http://www.rngr.net/Publications/ttsm
- Publisher
- USDA Forest Service.
- Year
- 2002
- ISBN
-
- Description
- An extremely good guide, it can also be downloaded in its entirety from the address shown above.
]. It has a round, sometimes open, crown with dense foliage and a bole that can be up to 1 metre in diameter[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
,
337- Title
- Tropical Tree Seed Manual
- Publication
-
- Author
- Vosso. J. Editor.
- Website
- http://www.rngr.net/Publications/ttsm
- Publisher
- USDA Forest Service.
- Year
- 2002
- ISBN
-
- Description
- An extremely good guide, it can also be downloaded in its entirety from the address shown above.
].
A valuable multi-purpose tree, yielding a valuable gum resin, a valuable timber and having a range of medicinal and other uses. The best form of the gum is generally considered to come from the var Myroxylon balsamum pereirae (see separate entry). This gum has a long history of use by local people as a medicine and is also widely used in modern herbal medicine, as an incense, perfume etc[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
,
318- Title
- Raintree Nutrition - Tropical Plant Database
- Publication
-
- Author
-
- Website
- http://www.rain-tree.com/plants.htm
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- Detailed information on the medicinal uses of more than two hundred rainforest plants, mainly from S. America, with basic descriptions of the plants, their other uses etc.
]. It is commonly gathered from the wild, but the tree is also sometimes cultivated for its gum, which is traded internationally[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. The tree is widely grown as an ornamental in the tropics, especially as a shade tree along the sides of roads[
].
Known Hazards
At least six clinical studies published in recent years indicate that balsam can cause allergic reactions in sensitive individuals. Allergic reactions reported are generally skin rashes and dermatitis when the balsam comes into contact with the skin - even in small amounts found in soaps, perfumes, and other common body care products[
318- Title
- Raintree Nutrition - Tropical Plant Database
- Publication
-
- Author
-
- Website
- http://www.rain-tree.com/plants.htm
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- Detailed information on the medicinal uses of more than two hundred rainforest plants, mainly from S. America, with basic descriptions of the plants, their other uses etc.
]. These allergic reactions are attributed to the gum's benzoic acids, which some people are highly sensitive to[
318- Title
- Raintree Nutrition - Tropical Plant Database
- Publication
-
- Author
-
- Website
- http://www.rain-tree.com/plants.htm
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- Detailed information on the medicinal uses of more than two hundred rainforest plants, mainly from S. America, with basic descriptions of the plants, their other uses etc.
].
Botanical References
Range
Northern S. America - western Brazil, Venezuela, Colombia and Ecuador, north through Central America to Mexico.
Habitat
An emergent tree in evergreen tropical humid forests in low to medium elevations from 100 to 600 metres[
337- Title
- Tropical Tree Seed Manual
- Publication
-
- Author
- Vosso. J. Editor.
- Website
- http://www.rngr.net/Publications/ttsm
- Publisher
- USDA Forest Service.
- Year
- 2002
- ISBN
-
- Description
- An extremely good guide, it can also be downloaded in its entirety from the address shown above.
].
Properties
Edibility Rating | |
Medicinal Rating | |
Other Uses Rating | |
Habit | Tree |
Height | 30.00 m |
Growth Rate | Slow |
Cultivation Status | Cultivated, Ornamental, Wild |
Cultivation Details
A plant of the moist to wet lowland tropics, where it is found at elevations up to 700 metres[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. It grows best in areas where annual daytime temperatures are within the range 22 - 28°c, but can tolerate 16 - 32°c[
]. When dormant, the plant can survive temperatures down to about -1°c[
]. It prefers a mean annual rainfall in the range 2,000 - 2,700mm, but tolerates 1,350 - 4,000mm[
].
Grows best in a sunny position, tolerating some shade[
]. Prefers a well-drained soil[
337- Title
- Tropical Tree Seed Manual
- Publication
-
- Author
- Vosso. J. Editor.
- Website
- http://www.rngr.net/Publications/ttsm
- Publisher
- USDA Forest Service.
- Year
- 2002
- ISBN
-
- Description
- An extremely good guide, it can also be downloaded in its entirety from the address shown above.
]. Plants can succeed on poor soils if they are well-drained[
]. Grows on mildly acid soils, preferring a pH in the range 5 - 8[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. Prefers a pH in the range 5.5 - 6.5, tolerating 5 - 8[
].
Seedlings grow to 2.5 metres tall within 2 years[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. Trees can attain a height of 10 metres in 10 - 12 years, and 20 metres in 25 years[
337- Title
- Tropical Tree Seed Manual
- Publication
-
- Author
- Vosso. J. Editor.
- Website
- http://www.rngr.net/Publications/ttsm
- Publisher
- USDA Forest Service.
- Year
- 2002
- ISBN
-
- Description
- An extremely good guide, it can also be downloaded in its entirety from the address shown above.
].
The trees are not a profitable source of balsam until about their 15th year[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. Under proper management, trees can yield gum for 30 - 40 years[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. Gum harvesting begins on 20 - 30 year old trees with a minimum diameter of 12 - 15 cm[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].
Trees commence flowering when about 5 years old[
337- Title
- Tropical Tree Seed Manual
- Publication
-
- Author
- Vosso. J. Editor.
- Website
- http://www.rngr.net/Publications/ttsm
- Publisher
- USDA Forest Service.
- Year
- 2002
- ISBN
-
- Description
- An extremely good guide, it can also be downloaded in its entirety from the address shown above.
].
Twenty year old trees yield about 3 kg of gum per year[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. Wild populations are the major sources of Peru and tolu balsam. Gum harvesting is often drastic and may cause serious tree injury or deaths[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].
There are various methods used to obtain the gum:-
1). Numerous V-shaped incisions are made in the bark, being careful not to girdle the tree[
413- Title
- Global Invasive Species Database
- Publication
-
- Author
-
- Website
- http://www.issg.org/database/welcome/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- Very detailed information on almost 400 species (with more being added) of plants that have become weeds in areas outside their native range.
]. The balsam flows through the incisions, into vases[
337- Title
- Tropical Tree Seed Manual
- Publication
-
- Author
- Vosso. J. Editor.
- Website
- http://www.rngr.net/Publications/ttsm
- Publisher
- USDA Forest Service.
- Year
- 2002
- ISBN
-
- Description
- An extremely good guide, it can also be downloaded in its entirety from the address shown above.
].
2). A section of bark 15 - 25 cm in size is removed from the tree at about 30 cm above the ground[
337- Title
- Tropical Tree Seed Manual
- Publication
-
- Author
- Vosso. J. Editor.
- Website
- http://www.rngr.net/Publications/ttsm
- Publisher
- USDA Forest Service.
- Year
- 2002
- ISBN
-
- Description
- An extremely good guide, it can also be downloaded in its entirety from the address shown above.
]. The wound is covered with a piece of cotton cloth to absorb the liquid. When the natural flow ceases, scorching stimulates it. The impregnated rags are then boiled in water and pressed[
337- Title
- Tropical Tree Seed Manual
- Publication
-
- Author
- Vosso. J. Editor.
- Website
- http://www.rngr.net/Publications/ttsm
- Publisher
- USDA Forest Service.
- Year
- 2002
- ISBN
-
- Description
- An extremely good guide, it can also be downloaded in its entirety from the address shown above.
].
3). Trees are burned at the base. Strips of bark are pulled off, crushed and placed in hot water to soften the balsam and facilitate its flow. The cooled balsam sinks to the bottom and can be separated[
413- Title
- Global Invasive Species Database
- Publication
-
- Author
-
- Website
- http://www.issg.org/database/welcome/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- Very detailed information on almost 400 species (with more being added) of plants that have become weeds in areas outside their native range.
].
4). Sections of the tree trunk are beaten with a wooden club and then vertical incisions 8 cm wide are made in the bark. A few days later the incisions are heated with fire to stimulate gum flow, but the incisions are not burned. Rags are placed over the incisions and removed when they are saturated. Crude presses are used to extract gum from the rags[
413- Title
- Global Invasive Species Database
- Publication
-
- Author
-
- Website
- http://www.issg.org/database/welcome/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- Very detailed information on almost 400 species (with more being added) of plants that have become weeds in areas outside their native range.
].
This species has a symbiotic relationship with certain soil bacteria, these bacteria form nodules on the roots and fix atmospheric nitrogen. Some of this nitrogen is utilized by the growing plant but some can also be used by other plants growing nearby[
755- Title
- Nodulation Plants in GRIN Taxonomy
- Publication
-
- Author
-
- Website
- http://www.ars-grin.gov/~sbmljw/cgi-bin/taxnodul.pl?language=en
- Publisher
- United States Department of Agriculture
- Year
- 0
- ISBN
-
- Description
- An online database listing plants that have either positive or negative reports on root and stem nodulation with nitrogen-fixing bacteria.
].
Edible Uses
The aromatic, slightly bitter gums obtained from this plant are used as a flavouring in soft drinks, confectionaries, ice cream and chewing gums[
301- Title
- Cornucopia II
- Publication
-
- Author
- Facciola. S.
- Publisher
- Kampong Publications, California.
- Year
- 1998
- ISBN
- 0-9628087-2-5
- Description
- The second edition of an excellent guide to the edible uses of plants, though it does not give any details of cultivation etc.
,
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].
The essential oil is used in flavouring baked goods. Its fragrance is attributed to vanillin, coumarin, cinnamic and benzoic acids[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].
Medicinal
The tree has a long history of use by local people, who used the leaves and fruit in the treatment of asthma, colds and flu, rheumatism, and external wounds[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. The sap is used for treating colds and lung ailments, abscesses, asthma, bronchitis, catarrh, headache, rheumatism, sores, sprains, tuberculosis, venereal diseases, and wounds[
318- Title
- Raintree Nutrition - Tropical Plant Database
- Publication
-
- Author
-
- Website
- http://www.rain-tree.com/plants.htm
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- Detailed information on the medicinal uses of more than two hundred rainforest plants, mainly from S. America, with basic descriptions of the plants, their other uses etc.
].
Many of these traditional uses have been borne out by modern research[
318- Title
- Raintree Nutrition - Tropical Plant Database
- Publication
-
- Author
-
- Website
- http://www.rain-tree.com/plants.htm
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- Detailed information on the medicinal uses of more than two hundred rainforest plants, mainly from S. America, with basic descriptions of the plants, their other uses etc.
]. The balsam of both varieties have been documented to have antiseptic, antiparasitic, and antibacterial properties as well as to promote the growth of epithelial (tissue) cells[
318- Title
- Raintree Nutrition - Tropical Plant Database
- Publication
-
- Author
-
- Website
- http://www.rain-tree.com/plants.htm
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- Detailed information on the medicinal uses of more than two hundred rainforest plants, mainly from S. America, with basic descriptions of the plants, their other uses etc.
].
In studies, the plants have been reported to inhibit Mycobacterium tuberculosis as well as the common ulcer-causing bacteria, H. Pylori in test-tube studies[
318- Title
- Raintree Nutrition - Tropical Plant Database
- Publication
-
- Author
-
- Website
- http://www.rain-tree.com/plants.htm
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- Detailed information on the medicinal uses of more than two hundred rainforest plants, mainly from S. America, with basic descriptions of the plants, their other uses etc.
].
The essential oil contains benzoic and cinnamic acid esters. These are believed to be the main active constituents of the balsam[
318- Title
- Raintree Nutrition - Tropical Plant Database
- Publication
-
- Author
-
- Website
- http://www.rain-tree.com/plants.htm
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- Detailed information on the medicinal uses of more than two hundred rainforest plants, mainly from S. America, with basic descriptions of the plants, their other uses etc.
].
Modern herbalism values tolu balsam, which is obtained from var balsamum, as a feeble expectorant used in cough mixtures and as an inhalant for catarrh and bronchitis[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. Balsam of tolu was included in the U.S. Pharmacopeia in 1820 and is used much as Peru balsam[
318- Title
- Raintree Nutrition - Tropical Plant Database
- Publication
-
- Author
-
- Website
- http://www.rain-tree.com/plants.htm
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- Detailed information on the medicinal uses of more than two hundred rainforest plants, mainly from S. America, with basic descriptions of the plants, their other uses etc.
]. In addition, it is a cough suppressant and respiratory aid used in cough lozenges and syrups, for sore throats, and as a vapour inhalant for respiratory ailments[
318- Title
- Raintree Nutrition - Tropical Plant Database
- Publication
-
- Author
-
- Website
- http://www.rain-tree.com/plants.htm
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- Detailed information on the medicinal uses of more than two hundred rainforest plants, mainly from S. America, with basic descriptions of the plants, their other uses etc.
].
Agroforestry Uses:
Tolu Balsam is a good shade tree and also fixes atmospheric nitrogen[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. It is sometimes grown to provide shade in coffee plantations[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].
Other Uses
'Tolu Balsam' exudes from injuries to the trunk and is obtained by cutting 'v'-shaped gashes in the bark. The fresh balsam is a soft, tenacious, yellowish-brown mass, but gradually hardens and eventually becomes brittle. It has a pleasant, fragrant odour and a balsamic taste. The balsam contains about 80% resin[
64- Title
- Vegetable Gums and Resins.
- Publication
-
- Author
- Howes. F. N.
- Publisher
- Faber
- Year
-
- ISBN
- 81-88818-17-8
- Description
- A very good book dealing with the subject in a readable way.
]..
The balsam is used in perfumery and for medicinal purposes. It has also been used as an adulterant for the more valuable 'Peru balsam' obtained from Myroxylon balsamum var pereirae[
64- Title
- Vegetable Gums and Resins.
- Publication
-
- Author
- Howes. F. N.
- Publisher
- Faber
- Year
-
- ISBN
- 81-88818-17-8
- Description
- A very good book dealing with the subject in a readable way.
].
The balsam extracted from the trunkt is used as an ingredient in commercial cosmetic preparations as a perfume[
].
The essential oil extracted from the balsmat obtained from the plant is used as an ingredient in commercial cosmetic preparations as a perfume[
].
The oleoresin extracted from the trunk is used as an ingredient in commercial cosmetic preparations as a film former, masking agent and hair conditioner[
].
Balsam gum contains about 60% cinnamein, an essential oil extracted by steam distillation[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. It is used as incense in churches and as a hair set and thickening agent[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. The oil is also used in high-grade perfume, cosmetic and soap industries[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. Its fragrance is attributed to vanillin, coumarin, cinnamic and benzoic acids[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].
The essential oil contains about 60% cinnamein, a volatile oil that is extracted by steam distillation and used commercially in the perfume, cosmetic, and soap industries[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].
The powdered bark is used as an underarm deodorant[
318- Title
- Raintree Nutrition - Tropical Plant Database
- Publication
-
- Author
-
- Website
- http://www.rain-tree.com/plants.htm
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- Detailed information on the medicinal uses of more than two hundred rainforest plants, mainly from S. America, with basic descriptions of the plants, their other uses etc.
].
The heartwood is reddish brown, turning deep red or purplish upon exposure, and has a spicy scent, it is sharply demarcated from the very pale yellow to white sapwood. The texture is medium to fine; the grain interlocked; lustre is medium; there is no distinct taste but a slightl distinctive aroma. The wood is moderately difficult to work, but can be finished smoothly with a high natural polish, and is very resistant to fungal decay. It is used for flooring, furniture, cabinetwork, turnery and railroad ties[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
,
316- Title
- Tropical Timbers of the World. Ag. Handbook No. 607.
- Publication
-
- Author
- Chudnoff. Martin.
- Publisher
- USDA Forest Service. Wisconsin.
- Year
- 1984
- ISBN
-
- Description
- Terse details on the properties of the wood of almost 400 species of trees from the Tropics.
,
363- Title
- Technical Information on 134 Species of Bolivian Woods
- Publication
-
- Author
- USDA Forest Service -- Forest Products Lab
- Website
- http://www2.fpl.fs.fed.us/publications/
- Publisher
- USDA Forest Service
- Year
- 0
- ISBN
-
- Description
- A PDF file, available for download. Exactly what the title says, brief and to the point.
]. The wood is not commercially marketed[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].
Propagation
Seed - it is mainly propagated through natural regeneration but seed can also be planted in a mixture of clay and organic matter to a depth of 5mm, covered with fine soil and watered daily[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. Germination beds or containers should be partially shaded[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. Seeds germinate in 15 - 30 days with a success rate of over 50%[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. Seedlings are ready for planting out in 5 months[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].
Natural regeneration is abundant mainly under the crown of parent trees, where seeds attain up to 80 percent germination[
337- Title
- Tropical Tree Seed Manual
- Publication
-
- Author
- Vosso. J. Editor.
- Website
- http://www.rngr.net/Publications/ttsm
- Publisher
- USDA Forest Service.
- Year
- 2002
- ISBN
-
- Description
- An extremely good guide, it can also be downloaded in its entirety from the address shown above.
]. Few of these seedlings ever reach maturity because of the low light levels in the forest[
337- Title
- Tropical Tree Seed Manual
- Publication
-
- Author
- Vosso. J. Editor.
- Website
- http://www.rngr.net/Publications/ttsm
- Publisher
- USDA Forest Service.
- Year
- 2002
- ISBN
-
- Description
- An extremely good guide, it can also be downloaded in its entirety from the address shown above.
].
Fruits must be soaked in running water for 24 hours in order to soften the pericarp and facilitate seed extraction[
337- Title
- Tropical Tree Seed Manual
- Publication
-
- Author
- Vosso. J. Editor.
- Website
- http://www.rngr.net/Publications/ttsm
- Publisher
- USDA Forest Service.
- Year
- 2002
- ISBN
-
- Description
- An extremely good guide, it can also be downloaded in its entirety from the address shown above.
].
The presence of high levels of coumarin in the cotyledons and embryonic axis seems to have no effect on the germination of this species, but it seems to act as an allelopathic factor[
303- Title
- World Agroforesty Centre
- Publication
-
- Author
-
- Website
- http://www.worldagroforestry.org/
- Publisher
-
- Year
- 0
- ISBN
-
- Description
- An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].