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Useful Tropical Plants

Phyllanthus emblica

L.

Phyllanthaceae

+ Synonyms

Cicca emblica (L.) Kurz

Diasperus emblica (L.) Kuntze

Dichelactina nodicaulis Hance

Emblica arborea Raf.

Emblica officinalis Gaertn.

Phyllanthus glomeratus Roxb. ex Benth.

Phyllanthus mairei H.Lév.

Phyllanthus mimosifolius Salisb.

Phyllanthus taxifolius D.Don

Common Name: Emblic

Phyllanthus emblica
Fruiting tree
Photograph by: Yosri
GNU Free Documentation License
Phyllanthus emblica Phyllanthus emblica Phyllanthus emblica Phyllanthus emblica

General Information

Emblic is a deciduous shrub or small tree; it usually grows 7.5 - 18 metres tall but occasionally can reach up to 30 metres. The bole, which is often crooked and gnarled, is up to 35cm in diameter. While actually deciduous, shedding its branchlets as well as its leaves, the plant is seldom entirely bare and is therefore often cited as being evergreen[
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
,
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
].
The plant is commonly cultivated in the home garden, especially in India, for its edible fruit and as a medicinal plant[
301
Title
Cornucopia II
Publication
 
Author
Facciola. S.
Publisher
Kampong Publications, California.
Year
1998
ISBN
0-9628087-2-5
Description
The second edition of an excellent guide to the edible uses of plants, though it does not give any details of cultivation etc.
] and is often found in local markets[
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
]. It is also grown as an ornamental[
200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.
], and is planted as a pioneer species in reforestation projects in Thailand[
998
Title
Forest Restoration Planting in Northern Thailand
Publication
Proceedings of the Southeast Asian Moving Workshop on Conservation, Management and Utilization of Forest Genetic Resources No. 31/2002
Author
Pakkad G.; Elliott S.; Anusarnsunthorn V. Et al
Website
http://www.fao.org/
Publisher
Food and Agriculture Organization of the United Nations; Bangkok
Year
2002
ISBN
 
Description
A report on reforestation in northern Thailand by using specific pioneer species. It can be downloaded from the Internet.
].

Known Hazards

None known

Botanical References

200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.
,
266
Title
Flora of China
Publication
 
Author
 
Website
http://flora.huh.harvard.edu/china/
Publisher
Missouri Botanical Garden Press; St. Louis.
Year
1994
ISBN
 
Description
An excellent, comprehensive resource in 25 volumes. In addition to the botanical information the flora also gives basic information on habitat and some uses. An on-line version is also available.

Range

E. Asia - China, Indian subcontinent, Myanmar, Thailand, Cambodia, Laos, Vietnam, Malaysia, Indonesia.

Habitat

Mixed forests[
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
]. Drier forests[
307
Title
Tropical and Subtropical Trees - A Worldwide Encyclopaedic Guide.
Publication
 
Author
Barwick. M.
Publisher
Thames & Hudson, London
Year
2004
ISBN
0-500-51181-0
Description
A superb book, very concise and well written, giving a wealth of information on 400 or more species including descriptions, habitat, cultivation details and plant uses. A wealth of colour photographs bring each plant vividly to life.
]. Dry open sparse forests or scrub, village groves at elevations of 200 - 2,300 metres in southern China[
266
Title
Flora of China
Publication
 
Author
 
Website
http://flora.huh.harvard.edu/china/
Publisher
Missouri Botanical Garden Press; St. Louis.
Year
1994
ISBN
 
Description
An excellent, comprehensive resource in 25 volumes. In addition to the botanical information the flora also gives basic information on habitat and some uses. An on-line version is also available.
].

Properties

Edibility Rating *  *  *
Medicinal Rating *  *  *  *  *
Other Uses Rating *  *  *
HabitDeciduous Shrub
Height15.00 m
Growth RateSlow
PollinatorsBees
Self-fertileNo
Cultivation StatusCultivated, Ornamental, Wild

Cultivation Details

A plant mainly of the hot, tropical lowlands, succeeding in both humid and semi-arid areas[
200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.
,
335
Title
Perennial Edible Fruits of the Tropics : an Inventory
Publication
 
Author
Martin, F. W.; Campbell, C. W.; Rubertbe, R. M.
Publisher
U.S. Dept. of Agriculture, Agricultural Research Service
Year
1987
ISBN
 
Description
A booklet giving terse details of almost 300 species of edible fruits, plus regional lists of Botanical and Common names for over 2,500 less well-known edible fruits of the tropics.
]. It can also be found at elevations up to 2,300 metres in southern China. It grows best in areas where annual daytime temperatures are within the range 20 - 29°c, but can tolerate 14 - 35°c[
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
]. It prefers a mean annual rainfall in the range 1,500 - 2,500mm, but tolerates 700 - 4,200mm[
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
].
Emblic is a very easily grown plant, reported to thrive in regions that are too dry and on soil that is too poor for most other fruit crops[
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
]. Requires a position in full sun or part day shade, but is undemanding as to soil requirements so long as it is well-drained[
200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.
,
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
]. It can even succeed on alkaline soils, though in a highly alkaline soil (pH 8.0) nutritional deficiencies are evident[
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
]. For maximum productivity, the tree requires deep soil ranging from sandy loam to clay, light or heavy, and slightly acidic to slightly alkaline[
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
]. Plants seem to grow equally well under both arid and humid conditions[
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
]. Prefers a pH in the range 6 - 8, tolerating 5 - 8[
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
].
The tree is rather slow-growing and usually only bears fruit when 6 - 8 years old[
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
]. Seedlings take about 10 years to come into bearing[
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
].
The plant only produces flowers at a daylength between 12 - 13.5 hours[
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
].
Annual fruit yields may be about 15 - 25 kilos per tree[
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
]. Mature plants can yield 200 kilos of fruit per year[
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
].
Ripe fruits may be retained for several months on the tree without significant loss of quality. Because of this, a long period is available for picking the fruits for consumption[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
].
The tree coppices well and pollards fairly well[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
]. Coppiced shoots grow particularly vigorously, and coppicing is considered the system most suitable for the production and collection of tanbark on a commercial scale[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
].
Usually plantations need much weeding because the thin crowns do not form a closed canopy[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
].
There are some named varieties[
301
Title
Cornucopia II
Publication
 
Author
Facciola. S.
Publisher
Kampong Publications, California.
Year
1998
ISBN
0-9628087-2-5
Description
The second edition of an excellent guide to the edible uses of plants, though it does not give any details of cultivation etc.
].
Emblic is fire resistant, and is one of the first trees to recover after a fire[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
,
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
].
Plants are usually monoecious, but occasional dioecious forms are found.

Edible Uses

Fruit - raw or cooked[
301
Title
Cornucopia II
Publication
 
Author
Facciola. S.
Publisher
Kampong Publications, California.
Year
1998
ISBN
0-9628087-2-5
Description
The second edition of an excellent guide to the edible uses of plants, though it does not give any details of cultivation etc.
,
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
]. An acid, rather astringent flavour, they are not often eaten raw unless accompanied with sugar, salt or chillies to moderate them[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
,
301
Title
Cornucopia II
Publication
 
Author
Facciola. S.
Publisher
Kampong Publications, California.
Year
1998
ISBN
0-9628087-2-5
Description
The second edition of an excellent guide to the edible uses of plants, though it does not give any details of cultivation etc.
]. The astringency can be removed by steeping the fruits in brine for a few days[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
]. The fruits are more commonly used to make jams, jellies, tarts, chutneys etc[
301
Title
Cornucopia II
Publication
 
Author
Facciola. S.
Publisher
Kampong Publications, California.
Year
1998
ISBN
0-9628087-2-5
Description
The second edition of an excellent guide to the edible uses of plants, though it does not give any details of cultivation etc.
]. The fruit is often used as a wayside nibble to quench the thirst[
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
]. Rich in pectin, the fruit is said to be one of the richest natural sources of vitamin C[
200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.
,
298
Title
Edible Leaves of the Tropics
Publication
 
Author
Martin. F. W.; Ruberte. R. M. & Meitzner. L. S.
Publisher
Echo. USA.
Year
1998
ISBN
0-9653360-1-8
Description
A terse guide, giving reasonable detail on the more common leaf crops and brief information on a wide range of lesser known plants.
,
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
], the fruit is also a good source of carbohydrate and minerals[
200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.
]. The mature fruit contains 1 - 1.8% vitamin C[
266
Title
Flora of China
Publication
 
Author
 
Website
http://flora.huh.harvard.edu/china/
Publisher
Missouri Botanical Garden Press; St. Louis.
Year
1994
ISBN
 
Description
An excellent, comprehensive resource in 25 volumes. In addition to the botanical information the flora also gives basic information on habitat and some uses. An on-line version is also available.
]. The yellow fruit is up to 25mm in diameter[
200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.
]. Fruits of wild plants weigh approximately 5.5g, cultivated fruits average 28 - 50g[
Many Hindus regard emblic as sacred and the Hindu religion prescribes that ripe fruit be eaten for 40 days after a fast in order to restore health and vitality. It is a common practice for Indian housewives to cook the fruits with sugar and saffron and give one or two to a child every morning[
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
].
Fruits are often preserved by splitting, removing the stone, putting the segments into a solution of 42% glycerol, 42% sucrose, water and preservatives, then heating to 90°c for 3 minutes. The fruits are allowed to equilibrate in the solution for two days at 2°c, then they are drained and packed into containers. Fruits preserved in this way remain acceptable for about 2 months at room temperature, and much longer when cooled, but the ascorbic acid content drops slowly[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
].

Leaves - cooked. Small, with a bitter flavour[
298
Title
Edible Leaves of the Tropics
Publication
 
Author
Martin. F. W.; Ruberte. R. M. & Meitzner. L. S.
Publisher
Echo. USA.
Year
1998
ISBN
0-9653360-1-8
Description
A terse guide, giving reasonable detail on the more common leaf crops and brief information on a wide range of lesser known plants.
].

Unripe seed[
301
Title
Cornucopia II
Publication
 
Author
Facciola. S.
Publisher
Kampong Publications, California.
Year
1998
ISBN
0-9628087-2-5
Description
The second edition of an excellent guide to the edible uses of plants, though it does not give any details of cultivation etc.
].
Seeds yield about 16% of a brownish yellow oil with linoleic acid (44%), oleic acid (28.4%), linolenic acid (8.8%), stearic acid (2.2%), palmitic acid (3.0%) and myristic acid (1.0%)[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
].

Used in the manufacture of black salt[
301
Title
Cornucopia II
Publication
 
Author
Facciola. S.
Publisher
Kampong Publications, California.
Year
1998
ISBN
0-9628087-2-5
Description
The second edition of an excellent guide to the edible uses of plants, though it does not give any details of cultivation etc.
].

Medicinal

Emblic is of great importance in traditional Asiatic medicine, not only as an antiscorbutic, but also in the treatment of diverse ailments, especially those associated with the digestive organs. In Thailand emblic fruits are traditionally used as an expectorant, antipyretic, diuretic, antidiarrhoeal and antiscorbutic[
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
]. Many of these traditional uses have been confirmed by research into the active ingredients in the plants and their properties.
The fruits, bark and leaves are rich in tannin[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
]. The dried pulp of unripe fruits contains 18 - 35% tannin; the content of ripe fruits is much lower. The dry stem bark contains 8 - 20% tannin. The bark of twigs is usually richer, containing 12 - 24% tannin on dry weight basis. Leaves may yield 22 - 28% tannin[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
].
The tannins of the fruit belong to the group of gallotannins and ellagitannins, giving on hydrolysis large amounts of gallic acid, small amounts of ellagic acid, and glucose[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
].
The tannin of the bark belongs to the group of proanthocyanidins, giving (+)leucodelphinidin on hydrolysis[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
].
The fruit is an extremely rich source of ascorbic acid, 100g of juice containing 600 - 1,300mg, sometimes even more[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
]. The tannin in the fruit prevents or retards the oxidation of the vitamin, so that the fruits can be preserved in salt solution or as dry powder while still maintaining their antiscorbutic value[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
].
The tannoid principles are potent inhibitors of aldose reductase and may be effective in the management of diabetic complications, including cataract[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
]
The fruits are a rich source of pectin[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
].
Many of the medicinal applications of the fruits can be ascribed to the presence of ascorbic acid and to the astringent action of the tannins, but the fruits contain other active compounds as well. Fruit extracts showed anti-oxidant and antitumour activities in in-vitro and animal tests. They also exhibited cholesterol-lowering, antitussive, anti-ulcerative and hepatoprotective properties and showed potent inhibitory activity on HIV reverse transcriptase; for the latter activity putranjivain A was the most active compound isolated[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
]
Phyllemblin has also been isolated from the fruits; it potentiates the action of adrenaline, has a mild depressant action on the central nervous system and has spasmolytic properties[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
].
Leaf extracts have shown inhibitory activity on human leukocytes and platelets, which at least partly confirms their anti-inflammatory and antipyretic properties[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
].

The fruits have diuretic, laxative and purgative activities and also show molluscicidal and antimicrobial properties. A principal ingredient of various Ayurvedic tonic formulae[
301
Title
Cornucopia II
Publication
 
Author
Facciola. S.
Publisher
Kampong Publications, California.
Year
1998
ISBN
0-9628087-2-5
Description
The second edition of an excellent guide to the edible uses of plants, though it does not give any details of cultivation etc.
], the fruit is given in order to allay the effects of aging and to restore the organs[
254
Title
The Encyclopedia of Medicinal Plants
Publication
 
Author
Chevallier. A.
Publisher
Dorling Kindersley. London
Year
1996
ISBN
9-780751-303148
Description
An excellent guide to over 500 of the more well known medicinal herbs from around the world.
]. The sour fruits are one of the ingredients of 'triphala', an Ayurvedic rejuvenating, laxative tonic based on this species plus the fruits of Terminalia bellirica and Terminalia chebula[
238
Title
Encyclopaedia of Herbs and their Uses.
Publication
 
Author
Bown. D.
Publisher
Dorling Kindersley, London.
Year
1995
ISBN
0-7513-020-31
Description
A very well presented and informative book on herbs from around the globe. Plenty in it for both the casual reader and the serious student. Just one main quibble is the silly way of having two separate entries for each plant.
]. The juice of the fruit is also given in order to strengthen the pancreas of diabetics, as well as in the treatment of eye problems, joint pain, diarrhoea and dysentery[
254
Title
The Encyclopedia of Medicinal Plants
Publication
 
Author
Chevallier. A.
Publisher
Dorling Kindersley. London
Year
1996
ISBN
9-780751-303148
Description
An excellent guide to over 500 of the more well known medicinal herbs from around the world.
].

Agroforestry Uses:

The branches are lopped for green manure[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
,
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
]. They are said to correct excessively alkaline soils[
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
].
The tree is planted as a pioneer species in northern Thailand in reforestation projects to restore native woodland - it is planted in degraded woodland and open areas in a mix with various other species that all have the ability to grow fast; produce dense, weed-suppressing crowns; and attract seed-dispersing wildlife, particularly birds and bats[
998
Title
Forest Restoration Planting in Northern Thailand
Publication
Proceedings of the Southeast Asian Moving Workshop on Conservation, Management and Utilization of Forest Genetic Resources No. 31/2002
Author
Pakkad G.; Elliott S.; Anusarnsunthorn V. Et al
Website
http://www.fao.org/
Publisher
Food and Agriculture Organization of the United Nations; Bangkok
Year
2002
ISBN
 
Description
A report on reforestation in northern Thailand by using specific pioneer species. It can be downloaded from the Internet.
].

Other Uses

The bark, as well as the roots, leaves and immature fruits, are highly valued as a source of tannins[
46
Title
Dictionary of Economic Plants.
Publication
 
Author
Uphof. J. C. Th.
Publisher
Weinheim
Year
1959
ISBN
-
Description
An excellent and very comprehensive guide but it only gives very short descriptions of the uses without any details of how to utilize the plants. Not for the casual reader.
,
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
,
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
,
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
].
The bark of shoots less than 5 cm in diameter is used to obtain a good tannin. Usually branches can be coppiced every 2 years. Quickly-dried bark contains much more tannin than slowly-dried bark. Therefore it has been recommended to dry the bark rapidly in the sun[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
].

The leaves are used for dyeing matting, bamboo wickerwork, silk and wool into brown colours. Grey and black colours are obtained when iron salts are used as mordants[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
].
Matting can be dyed dark colours with a decoction of the bark[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
].
The fruits are used to prepare a black ink and a hair dye[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
].

Branches and chips of the wood are thrown into muddy streams to clear the water and to impart a pleasant flavour[
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
,
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
].

The dried leaves are sometimes used as fillings in pillows[
266
Title
Flora of China
Publication
 
Author
 
Website
http://flora.huh.harvard.edu/china/
Publisher
Missouri Botanical Garden Press; St. Louis.
Year
1994
ISBN
 
Description
An excellent, comprehensive resource in 25 volumes. In addition to the botanical information the flora also gives basic information on habitat and some uses. An on-line version is also available.
].

The dried fruits are said to have detergent properties and are used for washing the head in parts of Nepal[
272
Title
Plants and People of Nepal
Publication
 
Author
Manandhar. N. P.
Publisher
Timber Press. Oregon.
Year
2002
ISBN
0-88192-527-6
Description
Excellent book, covering over 1,500 species of useful plants from Nepal together with information on the geography and peoples of Nepal. Good descriptions of the plants with terse notes on their uses.
].
A fixed oil derived from the fruit allegedly acts as a hair-restorer and is used in shampoos in India[
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].

A most curious custom is the making of simulated pottery jars from a paste of the boiled fruit, the surface being decorated with impressed coloured seeds[
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].

The red wood is close-grained, fairly heavy, hard but flexible, though highly subject to warping and splitting. It is used for minor construction, furniture, implements, gunstocks, hookahs and ordinary pipes[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
,
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
,
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
]. Very durable when submerged and believed to clarify water, it is utilized for making crude aqueducts and inner braces for wells[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
,
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
,
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
].
The wood is used as fuel and as a source of charcoal by villagers[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
,
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
]. It produces a good quality charcoal[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
].

Propagation

Seed - it does not store well and so is best sown as soon as it is ripe[
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. The seed is taken from over-ripe fruits, which are sun dried to facilitate removal of the stone, or are cut in half right through the stone. The extracted seeds are given the float test and 100% of those that sink will germinate. In 4 months, seedlings will have a stem diameter of 8 mm and can be budded or grafted if required[
306
Title
Under-Utilized Tropical Fruits of Thailand. RAP Publication 2001/26
Publication
 
Author
Suranant Subhadrabandhu
Publisher
F. A. O., The United Nations, Bangkok.
Year
2001
ISBN
 
Description
A detailed report on 35 fruit species that could have potential for increased production in Thailand.
].
Semi-hardwood cuttings, collected from the middle portions of vigorous shoots of young trees and planted in beds at a temperature of about 33°c, produce a high percentage of rooting, up to 84%[
299
Title
Protabase - Plant Resources of Tropical Africa.
Publication
 
Author
 
Website
http://www.prota.org
Publisher
 
Year
0
ISBN
 
Description
An excellent on-line database with detailed information on over 3,200 species of useful plants of Africa.
].
Greenwood cuttings.
Layering.
Grafting.

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