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Useful Tropical Plants

Bertholletia excelsa

Bonpl.

Lecythidaceae

+ Synonyms

Barthollesia excelsa Silva Manso

Bertholletia nobilis Miers

Common Name: Brazil Nut

Bertholletia excelsa
Tree
Photograph by: Nando cunha
Creative Commons Attribution 3.0
Bertholletia excelsa Bertholletia excelsa Bertholletia excelsa Bertholletia excelsa Bertholletia excelsa Bertholletia excelsa Bertholletia excelsa Bertholletia excelsa

General Information

Brazil nut is a large evergreen tree with a globose crown, that can grow 25 - 56 metres tall. The straight, cylindrical bole can be 1 - 2 metres in diameter and is unbranched until near the top[
200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.
,
454
Title
Useful Fiber Plants of the World
Publication
 
Author
Dodge C.R.
Website
http://www.biodiversitylibrary.org/
Publisher
USDA; Washington.
Year
1897
ISBN
 
Description
A rather dated, but very comprehensive catalogue of fibre plants from around the world. The book can be downloaded from the Internet.
].
The seed is one of the most commonly consumed nuts in the world. The tree is not usually cultivated, the seed being gathered in large quantities from the wild and then exported from there to many other countries. The tree is also harvested from the wild for its wood and for local medicinal use. The tree has not as yet been much cultivated since it is abundant in the Amazon jungle, though some work on cultivating it has been carried out successfully in other humid tropical areas such as Malaya, Sri Lanka and the West Indies[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
].

Known Hazards

Nuts from trees growing on barium-rich soils can accumulate up to 0.29% barium and should be avoided due to danger of barium toxicity[
324
Title
Edible Nuts.
Publication
Non-Wood Forest Products, Handbook 5
Author
Wickens G.E.
Website
http://www.fao.org/docrep/
Publisher
FAO, Rome.
Year
1995
ISBN
92-5-103748-5
Description
Detailed information on 20 of the most valuable nut crops, plus more basic information on a wide range of other nut-producing plants. It can be downloaded from the Internet.
].

Botanical References

200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.

Range

North and western S. America - Brazil, Venezuela, the Guyanas.

Habitat

Deep, rich, alluvial soils, on slightly raised ground that is not subject to regular or extensive flooding[
200
Title
The New RHS Dictionary of Gardening. 1992.
Publication
 
Author
Huxley. A.
Publisher
MacMillan Press
Year
1992
ISBN
0-333-47494-5
Description
Excellent and very comprehensive, though it contains a number of silly mistakes. Readable yet also very detailed.
].

Properties

Edibility Rating *  *  *  *  *
Medicinal Rating *  *
Other Uses Rating *  *  *
HabitEvergreen Tree
Height40.00 m
Growth RateMedium
PollinatorsBees
Self-fertileNo
Cultivation StatusCultivated, Wild

Cultivation Details

A plant of the lowland, tropical rain forests, where it is found at elevations up to 200 metres[
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
,
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
]. It grows best in areas where annual daytime temperatures are within the range 20 - 36°c, but can tolerate 12 - 40°c[
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
]. It prefers a mean annual rainfall in the range 2,400 - 2,800mm, but tolerates 1,400 - 3,500mm[
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
]. In eastern Amazonia, in the lower limits of its climatic range, there can be 2 - 7 months where the monthly rainfall is less than 100 mm[
324
Title
Edible Nuts.
Publication
Non-Wood Forest Products, Handbook 5
Author
Wickens G.E.
Website
http://www.fao.org/docrep/
Publisher
FAO, Rome.
Year
1995
ISBN
92-5-103748-5
Description
Detailed information on 20 of the most valuable nut crops, plus more basic information on a wide range of other nut-producing plants. It can be downloaded from the Internet.
]. A pronounced dry season is necessary for good fruit set[
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].
Requires a deep, well-drained but moisture retentive fertile, medium to heavy soil[
307
Title
Tropical and Subtropical Trees - A Worldwide Encyclopaedic Guide.
Publication
 
Author
Barwick. M.
Publisher
Thames & Hudson, London
Year
2004
ISBN
0-500-51181-0
Description
A superb book, very concise and well written, giving a wealth of information on 400 or more species including descriptions, habitat, cultivation details and plant uses. A wealth of colour photographs bring each plant vividly to life.
,
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
]. Mature trees grow best in a sunny position[
307
Title
Tropical and Subtropical Trees - A Worldwide Encyclopaedic Guide.
Publication
 
Author
Barwick. M.
Publisher
Thames & Hudson, London
Year
2004
ISBN
0-500-51181-0
Description
A superb book, very concise and well written, giving a wealth of information on 400 or more species including descriptions, habitat, cultivation details and plant uses. A wealth of colour photographs bring each plant vividly to life.
], but young trees require shade for their first 5 - 7 years[
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
]. Well adapted to heavy clay and low-fertility Oxisols but is not tolerant to waterlogging[
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. Prefers a pH in the range 5.5 - 7, tolerating 4.5 - 8[
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
].
Trees commence flowering when about 5 - 6 years old[
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
]. Fruiting starts at 12 - 16 years in the forest and as early as 8 years if trees are well managed in the open[
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
,
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
]. Mature nuts are produced approximately 15 months after fertilization[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
,
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].
The seeds are borne in a large spherical wooden 'fruit' that is up to 15cm in diameter and can weight up to 2 kilos[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
]. Between 12 and 24 seeds are contained in each fruit and this falls intact from the tree when it is ripe[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
]. Since the fruits are only formed on the upper branches, and these can be 30 metres or more above ground level, this can make collecting the seeds a rather hazardous occupation[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
].
The nuts are normally extracted in situ from the 'fruits' and then shipped out of the jungle to be sold[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
].
Trees 16 years old produce 30 - 50 fruits, mature trees usually 200 - 400, and a yield of 1,000 fruits has been reported[
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. Yields of up to 450 kilos of nuts can be obtained from the best trees[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
].

Edible Uses

Seed - raw or cooked. A mild flavour, the nuts are a common item of food, eaten raw and also roasted, salted or used in ice cream etc[
301
Title
Cornucopia II
Publication
 
Author
Facciola. S.
Publisher
Kampong Publications, California.
Year
1998
ISBN
0-9628087-2-5
Description
The second edition of an excellent guide to the edible uses of plants, though it does not give any details of cultivation etc.
]. The nuts are very nutritious, containing roughly 66% fat or oil, 17% protein and 7% carbohydrate[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
]. The ripe fruit is a wooden capsule weighing up to 750g and containing 12 - 24 angular brown seed, each about 5cm long and 2.5cm wide[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
,
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
].

An oil is obtained from the seed[
301
Title
Cornucopia II
Publication
 
Author
Facciola. S.
Publisher
Kampong Publications, California.
Year
1998
ISBN
0-9628087-2-5
Description
The second edition of an excellent guide to the edible uses of plants, though it does not give any details of cultivation etc.
]. The fresh-pressed oil is pale yellow, almost odourless, with a pleasant nutty flavour[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
]. It is not unlike almond oil in composition[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
]. It can be used like olive oil[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
].

Medicinal

The bark of the tree is used as a treatment for liver problems[
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].

The sweet oil from the seeds is applied to burns[
348
Title
Medicinal Plants of the Guianas
Publication
 
Author
DeFilipps, R. A.; Maina, S. L.; & Crepin, J.
Website
http://botany.si.edu/bdg/medicinal/index.html
Publisher
Smithsonian Museum
Year
0
ISBN
 
Description
A down-loadable PDF document of a book in pre-publication awaiting illustration. An excellent, if rather terse, guide to the traditional medicinal uses of the plants of the region
].

A tea made from the fruit is used as a treatment for gastralgia[
739
Title
Medicinal Plants of Brazil
Publication
 
Author
Mors W.B.; Rizzini C.T.; Pereira N.A.
Publisher
Reference Publications; Michigan.
Year
2000
ISBN
0-917256-42-5
Description
An extremely concise book, listing approximately 1,500 species and varieties of plants with their botanical name, common name in Brazil, regions in Brazil where the plant is found, a brief list of uses and any medically active compounds in the plant.
].

Other Uses

The hard woody 'fruit' that contains the seeds is used as cups or other receptacles[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
]. It is also made into bracelets or carved to make other ornaments[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
,
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].
Empty fruit capsules are used to carry small, smoky fires in order to discourage black flies (Simulium spp.) from attacking people working in the field during the rainy season[
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].
Open capsules are sometimes used to collect latex from rubber trees[
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].

The oil obtained from the seed has been used as an illuminant[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
]. The oil has also sometimes been used in perfumery, paint and soap making, and as a lubricant for watches and clocks[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
,
418
Title
Ecocrop
Publication
 
Author
 
Website
http://ecocrop.fao.org/ecocrop/srv/en/home
Publisher
 
Year
0
ISBN
 
Description
Basic information on a wide range of useful plants, plus details of environmental needs where available.
].

The woody 'fruits' make an excellent fuel[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
,
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. The nut, which has an elevated oil content of 63-69%, burns with a candle-like flame when lit[
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].

The bark has been used in remote regions for fibre production[
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. It is used for caulking ships[
46
Title
Dictionary of Economic Plants.
Publication
 
Author
Uphof. J. C. Th.
Publisher
Weinheim
Year
1959
ISBN
-
Description
An excellent and very comprehensive guide but it only gives very short descriptions of the uses without any details of how to utilize the plants. Not for the casual reader.
,
454
Title
Useful Fiber Plants of the World
Publication
 
Author
Dodge C.R.
Website
http://www.biodiversitylibrary.org/
Publisher
USDA; Washington.
Year
1897
ISBN
 
Description
A rather dated, but very comprehensive catalogue of fibre plants from around the world. The book can be downloaded from the Internet.
].

The heartwood is light brown to reddish; it is not clearly demarcated from the 3 - 5cm wide band of sapwood. The texture is medium, the grain straight or interlocked; lustre is medium. The wood is moderately heavy, hard and moderately durable being moderately resistant to fungi and termites but susceptible to dry wood borers. Seasoning is fairly fast with only a slight risk of checking or distortion; once dry it is moderately stable in service. It works well with normal tools; nailing and screwing are good; gluing is correct for internal purposes only. The durable wood is sought by boat builders, and is also used for purposes such as interior joinery, cabinet making, high class furniture, construction, flooring, heavy carpentry and sliced veneer[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
,
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
,
363
Title
Technical Information on 134 Species of Bolivian Woods
Publication
 
Author
USDA Forest Service -- Forest Products Lab
Website
http://www2.fpl.fs.fed.us/publications/
Publisher
USDA Forest Service
Year
0
ISBN
 
Description
A PDF file, available for download. Exactly what the title says, brief and to the point.
,
419
Title
Brazilian Trees. Volume 1. 4th Edition.
Publication
 
Author
Lorenzi. H.
Publisher
Instituto Plantarum De Estudos Da Flora; Brazil.
Year
2002
ISBN
85-86714-17-8
Description
Information on 350 species of Brazilian trees. Stunning photographs of each species, brief details on the plant, its uses and how to grow it from seed. A superb work, with the minor irritation that the translation from Portuguese is not of the best.
,
848
Title
Tropix 7
Publication
 
Author
 
Website
http://tropix.cirad.fr/en
Publisher
CIRAD
Year
0
ISBN
 
Description
An on-line guide to the timbers of 245 species of trees.
].
The felling of the trees is discouraged because they are more valuable as a nut crop, however the wood is well suited to all manner of structural work[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
].

Propagation

Seed - fresh seed germinates rapidly, often within 2 - 3 weeks, so long as a portion of the hard shell is cracked or carefully removed first in order to allow the entry of moisture[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
]. Otherwise seed can take 2 years to germinate[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
]. Seedlings probably need good light to grow away well[
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
]. Seedling trees have borne a first crop at an age of 14 - 20 years[
63
Title
Nuts.
Publication
 
Author
Howes. F. N.
Publisher
Faber
Year
1948
ISBN
-
Description
Rather old but still a masterpiece. Has sections on tropical and temperate plants with edible nuts plus a section on nut plants in Britain. Very readable.
].
If the conditions are right, some of the seeds buried by rodents may germinate. Trees appear to depend on natural or artificial disturbance to reproduce. For example, they will be found in clearings in the forest created by humans, forest gaps created by tree fall, or sprouting from damaged mother roots. Trees take about 10 years to start reproducing; grafted material may begin to bear nuts within 4 years. Other than using a single clone, several selections are cloned and grafted onto stock to encourage cross-fertilization, which increases yield[
303
Title
World Agroforesty Centre
Publication
 
Author
 
Website
http://www.worldagroforestry.org/
Publisher
 
Year
0
ISBN
 
Description
An excellent online database of a huge range of trees giving very good information on each plant - its uses, ecology, identity, propagation, pests etc.
].
Cite as: Tropical Plants Database, Ken Fern. tropical.theferns.info. 2019-09-21. <tropical.theferns.info/viewtropical.php?id=Bertholletia+excelsa>

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